In Toho’s movies, Godzilla has a deep rogues’ gallery of villains. Beginning in the 1960s, the King of the Monsters has had to fight off over a dozen villainous kaiju. Some have become popular enough to be appear in multiple movies, while others have only been used once.

Godzilla has been fighting with other monsters since 1955’s Godzilla Raids Again when he went up against Anguirus. But it’s important to keep in mind that not all of Godzilla’s foes are villains, per se. There are occasions when it’s Godzilla who’s the real antagonist. That’s why creatures like Mothra and Baragon can’t be counted as Godzilla villains, even though they’ve both fought him.

Related: Mothra Makes Godzilla's Weirdest Monster Possible For The MonsterVerse

Also, not all Godzilla movies are about good versus evil. There are times when he gets into brief fights with other monsters, who later become allies. Examples of this include Anguirus, Rodan, and Kumonga. On the other hand, there’s a multitude of creatures in the franchise who can only be classified as enemies of Godzilla. Here’s every Godzilla villain in Toho’s kaiju library.

King Ghidorah

King Ghidorah

The first time Godzilla battled a true villain on the big screen was in 1964’s Ghidorah, the Three-Headed Monster. Introduced as an alien creature sent to conquer the world, Ghidorah was driven off by the combined efforts of Godzilla and Rodan, who received some assistance from Mothra. Ghidorah returned a few more times in the Showa series, but was always defeated by Godzilla and at least one of his allies. It wasn’t until 1991’s Godzilla vs. King Ghidorah that the two monsters were able to have a fair, one-on-one fight. In Godzilla: Final Wars, he was completely reimagined as Monster X, who transformed into Keizer Ghidorah. Commonly regarded as Godzilla’s greatest villain ever, Ghidorah has fought Godzilla more than any other villain, with his most recent appearance being in the MonsterVerse’s Godzilla: King of the Monsters.

Kamacuras

Kamacuras

In Son of Godzilla, Godzilla had to defend Minila from three giant, preying mantises called Kamacuras. Godzilla killed two of them, while the third was disposed of by Kumonga, the giant spider. Godzilla had to fight another in Godzilla: Final Wars.

Ebirah

Ebirah

In the low-budget Ebirah, Horror of the Deep, Godzilla battled Ebirah, a giant crab monster who lived in the ocean and attacked any ships that drew near it. In combat, the creature relied mostly on its claws, and wasn’t able to do much against Godzilla once they were ripped off. It’s been said that Toho’s original intention for the movie was for Ebirah to fight King Kong, not Godzilla.

Related: KOTM May Have Secretly Set Up Godzilla's Most Important Human Character

Gabara

Gabara

Gabara, a giant green monster, stands out as a Toho kaiju that doesn’t actually exist in Godzilla canon. That’s because his only role was in All Monsters Attack, a movie that centers on a young boy’s dreams of adventuring on Monster Island with Godzilla’s son. Gabara was a manifestation of the boy’s fears about a bully at his school. The boy dealt with this fear by dreaming about Gabara being defeated by Godzilla.

Hedorah

Godzilla Fights Hedorah

Hedorah, also known as the Smog Monster, was an alien blob that fed on pollution. The threat that Hedorah represented was a direct result of humanity’s mistreatment of the Earth. He posed a unique challenge to the King of the Monsters in Godzilla vs. Hedorah. Godzilla had trouble beating him, as the toxic chemicals in his body did significant damage to his skin. Plus, Hedorah held up well against all of Godzilla’s attacks, including his atomic breath.

Gigan

Gigan

Gigan, a cyborg alien with hooks for hands, became the first monster to make Godzilla bleed in Godzilla vs. Gigan. In fact, in that movie he gave Godzilla what was arguably the most gruesome beating he’s ever received, which is a big part of why he’s considered one of Godzilla’s greatest enemies. That being said, he’s never been a singular antagonist in a Toho movie. He’s always an ally of another monster, such as Ghidorah and Megalon.

Megalon

Megalon

In Godzilla vs. Megalon, Godzilla and Jet Jaguar teamed up for an exciting showdown with the duo of Gigan and Megalon. Megalon was a giant cockroach-like monster who could burrow underground with his hands, which functioned like drills.

Related: Why Godzilla Should Beat Mechagodzilla Without Kong's Help

Mechagodzilla

Mechagodzilla

As his name implies, Mechagodzilla is the robotic counterpart of the King of the Monsters. Armed with a deadly arsenal of missiles and laser cannons, the mechanical kaiju is one of the most dangerous enemies Godzilla has ever faced. In Godzilla vs. Mechagodzilla and Terror of Mechagodzilla, he was portrayed as a villain created by aliens to help them take over the Earth. In each of his appearances that followed, Mechagodzilla has been utilized as a weapon of humanity meant to protect them from Godzilla.

Titanosaurus

Terror of Mechagodzilla giant monster fight

Unlike most Godzilla villains, Terror of Mechagodzilla’s Titanosaurus wasn’t an evil beast bent on destruction. In the beginning of the movie, he was described as a peaceful, aquatic creature, but after being mind-controlled by aliens, Titanosaurus joined forces with Mechagodzilla and helped him oppose Godzilla. Despite the odds, Godzilla managed to survive against both monsters. After his defeat, the creature never returned to the Godzilla universe.

Biollante

In Godzilla vs. Biollante, a terrifying new villain was born from the DNA of a rose, a dead girl, and Godzilla. At the time, Biollante was the largest monster he had ever encountered. The creature — who dwarfed even Godzilla himself — was an enormous plant monster whose long, vine-like tendrils could be used to subdue her enemies.

SpaceGodzilla

SpaceGodzilla

Powered by the large crystals on his shoulders, SpaceGodzilla is an evil kaiju who was created when Godzilla’s cells were mutated by the energy from a black hole. After coming to Earth, SpaceGodzilla captured Godzilla’s son and became embroiled in a heated battle with the Japanese military, a robot called Moguera, and of course, the monster he originated from. Due to the vast amount of power he carried, SpaceGodzilla is the only Toho villain that Godzilla was never able to beat without help.

Related: MonsterVerse Theory: Kong Kills Godzilla's Minions (So They Fight)

Battra

Battra

Godzilla vs. Mothra saw the first and only appearance of Battra, Mothra’s evil counterpart. It was explained in the movie that while Mothra is a protector of humanity, Battra hates humans and sees them as a threat to the planet. After battling Godzilla, Battra realized that he couldn’t win without forming an alliance with Mothra. In the end, Battra helped Mothra take down Godzilla, but lost his life in the process.

Destoroyah

Godzilla vs. Destoroyah, the final installment in the Heisei series, revealed that there was a side-effect of the military using the Oxygen Destroyer to kill the 1954 Godzilla. Apparently, the Oxygen Destroyer mutated a group of crustaceans into Destoroyah, an incredibly powerful and destructive being. Godzilla, who was leaking radiation and near the end of his life, barely managed to finish off Destoroyah before using what was left of his life-force to save his son.

Orga

Orga - Godzilla 2000

In the first film in the Millennium series, Godzilla 2000, the titular monster crossed paths with what appeared to be an alien spaceship. After absorbing some of Godzilla’s DNA, the spaceship morphed into Orga, a fearsome, hulking creature. Orga attempted to become a clone of Godzilla, but was destroyed before the transformation was complete.

Megaguirus

Like Orga, Destoroyah, Battra, and a few others, Megaguirus was a one-and-done villain. In Godzilla vs. Megagurius, a swarm of prehistoric dragonflies stole some of Godzilla’s power, and gave it their queen. Thanks to Godzilla’s energy, the dragonfly grew and became Megaguirus.

More: Godzilla: All 12 Monster Allies (& Which Movies They Appeared In)